Leganes has made club history by reaching the quarterfinals of the Copa del Rey for the first time, having shocked Villarreal on Wednesday.
A goal from Nordin Amrabat had given Leganes a slender advantage from the home leg of their last-16 tie.
And when Nabil El Zhar put his side ahead after half an hour at the Estadio de la Ceramica, Leganes was on track to for its maiden last-eight appearance.
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Villarreal battled back after the break, though, Dani Raba on target shortly after the interval to set up a grandstand finish.
And, even though Denis Cheryshev scored in the last minute to level the scores on aggregate, Leganes progressed to the quarterfinals on away goals.
Alaves are also in the hat after easing past Formentera, while Atletico Madrid comfortably dispatched of Lleida Esportiu 7-0 on aggregate on Tuesday.
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